The animals are some of the largest remaining megafauna, with all species able to reach or exceed one tonne in weight. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm) protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species lack teeth at the front of their mouths, relying instead on their lips to pluck food.^[1]